After twenty years of being away, Harlow has come back to his birth pack. Now in his early forties, he’s a doctor and ready to retire from human life in order to take his place in the pack. But the pack he left behind isn’t anything like what he comes back to. His older brother is the new alpha, and the pack that was once full of families is now little more than a harem of young wolves following his brother blindly. The only one not afraid of his brother is his beta, Nathan, who stays to protect the others. Harlow brings with him the much needed hope Nathan has been searching for, but Harlow has to prove himself worthy of the position of alpha before Nathan will follow him.

Jacob wants nothing to do with his father, but he's not fitting in with his birth pack anymore. When his father sends for him, Jacob's pack urges him to go, giving him little choice in the matter. Colorado is a far cry from Boston, and he just wants to find somewhere he fits in, wherever that may be. Miles is finally settling into life with a good alpha at the helm. He's learning what it means to be an omega and that he has a definite place in the pack. Jacob is a complication in ways he doesn't understand. He wants to be close to Jacob, but he's also afraid of what that means.

Gabriel isn't trying to constantly screw up in the Glenwood Springs pack, but it seems like no matter what he does, everything turns out to be his fault. When he lies to his alpha in order to go into town and hook up with a human, things go from bad to worse. Harlow has had enough and sends him off to a nearby pack to join their ranks. He has a cousin there, so he's not completely alone, but being rejected by two packs in less than six months is a hard blow to his over-inflated ego. Chris has heard a lot about Gabriel before he even gets to the pack, and none of it is good. While Chris is determined to be respectful because his best friend is Gabriel's cousin, that's as far as he's willing to go, and his patience is quickly wearing thin with each of Gabriel's crass comments. When Chris has enough and hits his breaking point, it could ruin Gabriel's chances of ever finding his place in a pack, or it could end up being the push Gabriel needs to learn how to belong. Regardless of what happens, Gabriel’s life is going to change.

Misha has never been the same since his mate and child died in a house fire. He was managing, but his thin hold on reality quickly frayed. As a feral wolf, he was lucky to find a pack willing to work with him and a beta available to watch over him. Xavier just wants Misha to be better. He's heard of the awful things that can turn wolves feral and also what most packs do to them. He's glad Misha didn't end up that way, and he wants to be able to help him, no matter what Misha needs from him. The pack is waiting for him to come around, and they're all ready to welcome Misha to their ranks, but it's up to him to come back from the dark thing he's turned into.

Some things are more important than duty. For years, Dean always did as his alpha instructed, it’s what pack enforcers were expected to do. When he’s ordered to track down a rogue who stole a powerful magic item and killed members of his old pack, he embarks on a dangerous mission without hesitation, even if it means entering neighboring territories without permission. Evan has endured years of trauma, and he’s not sure he’s ready to make himself vulnerable again. Even with a real alpha, and a supportive pack, the world is a scary place and he still bears the emotional scars of years of torture at the hands of someone who was supposed to take care of him. As they face the danger the rogue brings to both of them and the Woodlands pack, can their hearts open enough to help Evan mend what was torn from him? Will Evan help Dean find a new path he’d never expected? When they confront unknown magic, will it help, or inflict even more pain on them both? Pack Rules continues with more adventures of family and healing.