Meet the All-American pups of Buxton, U.S.A. Jake, Sheena, Fritz, Rosie and Tracker. They're cute. They're cuddly. They're frisky. And they just can't stay out of trouble! Jake, Sheena, Tracker, and Fritz are on the trail of their new friend, a puppy named Rosie, who has headed off to the big, scary city to find her lost owner. Will the puppies find Rosie before the dog pound catches them all?

Sheena loves playing with her puppy pals—chasing water rats and eating croissants at the Main Street Bakery. But now she has to spend her mornings with Amber, a prissy poodle who doesn't seem to know a thing about being a real puppy. Is Sheena doomed to spend her days sleeping on couches and having her nails painted with this sissy little fuzzball forever?

The top-secret cake recipe for the annual bake-off has been stolen! Tracker is on the case, using his nose as his guide. He leads his puppy friends on a wild chase, sniffing all the way. The pups are soon hot on the trail of the missing recipe. Will they catch the burglar before he steals first prize at the bake-off, too?