Nineteen-year-old Orsay lives an uneventful life in the French countryside, until the day he gains extraordinary powers in his hands after an encounter with a mysterious creature known as a whol. On a trip to Paris in search of a cure, he meets and falls for Basma, a passionate activist for whols' rights. But Orsay isn't convinced that whols should be considered equal to humans. Especially once Melek, another human with the same powers, embarks on a murderous rampage to avenge those she sees as her kin.

Not only are the mysterious creatures known as whols beautiful, a handful of humans who have acquired some of their powers can meld with them to enter a mind-bending space full of love, connection, and danger. Juliette's new powers in her eyes may hold the key to unlocking even more possibilities, if she can learn to accept herself... and if she, Orsay, and the rest of their friends can stay one step ahead of a strange woman bent on feeing their "whol parts" to her young son.

Après les évènements d'Étretat, Orsay, Basma et Juliette ont été amnistiés par la police, contrairement à Melek et Vicento, toujours recherchés. Le trio retourne chez les parents d'Orsay et cherchent par tous les moyens à soigner le cancer de la mère du jeune homme, en vain. Pendant ce temps, les gouvernements mondiaux tolèrent de moins en moins les toutes qui finissent par devenir hostiles. Les populations locales, cherchant d'abord à se défendre, se mettent alors à les détruire. Ne pouvant laisser un tel massacre se perpétrer, Orsay décide de prendre les choses en main et qui sait, trouver la « clé » qui pourrait soigner sa mère. Et le monde?